Proprioreception is the sense of position and movement.
There are three types of mechanoreceptors in muscles and joints:
Stretch-sensitive receptors in the skin, including Ruffini endings and Merkel discs, also signal postural information.
Somatotopic information is retained as nerves enter the spinal cord through dorsal roots.
diameter | velocity | myelinated | ||
Ia | 12-20 | 70-120 | yes | muscle spindle stretch - proprioreception |
Ib | 12-20 | 70-120 | yes | golgi tendon organs (contraction) |
II (Aβ) | 6-12 | 30-70 | yes | Meissner corpuscles, Merkel cells |
III (Aδ) | 1-5 | 5-30 | yes | free nerve endings: sharp pain, cold, hair receptors |
IV (C) | 1 | 0.5-2 | no | free nerve endings: dull pain, warmth, itch |
Ia fibres participate in spinal reflexes
There are many ways for sensory information to reach the cerebellum, both directly and through brainstem relays.
The posterior spinocerebellar tract carries information from muscle spindles and Golgi tendon organs, synapsing on the thoracic nucleus at the base of the dorsal horn. It ascend ipsilaterally and travels through the inferior cerebellar peduncle to reach the cerebellar vermis and adjoining areas. It primarily carries proprioreceptive information about the lower limbs. The cuneocerebellar tract travels alongside the PSCT and carries information about the arm.
The anterior spinocerebellar tract carries information from proprioreceptors, cutaneous receptors, spinal interneurons, and descending tract fibres. This integration allows it to carry complex information related to attempted movements. The ASCT crosses in the spinal cord and ascends to the superior pons, entering the cerebellum through the superior peduncle. It then recrosses the midline, ending in the ipsilateral vermis and adjoining areas.
The dorsal column and medial lemniscus also carry proprioreceptive information.